"I need to hear some sounds that recognize the pain in me, yeah. but the airways are clean and there's nobody singing to me now." The Verve, Bittersweet Symphony |
"Nascerá dentro me sul silenzio che habita qui. Fiorirá un canto che mai nessuno ha cantato per te." Gen Rosso, Nascerá |
"There is nothing in which the birds differ more from man
than the way in which they can
build and yet leave a landscape as it was before. "
Robert Lynd (1879 - 1949), The Blue Lion
Jamaican oriole (nominate race, Icterus leucopteryx leucopteryx). Anchovy, north-western Jamaica.
Hymns to Him in the Highest
The passerine birds are such a huge assemblage of birds that I have decided to discuss them in a section of their own.
ORDER PASSERIFORMES
The largest avian order is that of the songbirds. Including almost three quarters of modern dinosaurs on Earth, it is the most varied and successful group of living creatures with feathers. From tiny kinglets a few centimeters long to the huge common and Ethiopian ravens, songbirds occur in every shape and color, with every lifestyle, in every continent save for Antarctica, in every habitat except for the open oceans of the World. Some, like the Saint Andrew's vireo of the West Indies, are limited in their distribution, inhabiting tiny islands. Others, like the North American bobolink and the Arctic tern, are among the greatest migrants, whose travels extend among continents. Seed, fruit, insect, and carrion eaters, as well as hunters of lizards, snakes, mammals, and other birds; arboreal or terrestrial; from sea level to high alpine summits; from deserts to rain forests; with plumages ranging from somber blacks, grays, and browns to others that pass for living rainbows; songbirds of one kind or another can be found almost everywhere on land where man looks on land.
Within the passeriformes there are many different families. Some of them are discussed below.
Family Sylviidae: Gnatcatchers and Their Kin
Gnatcathchers are small birds, usually of drab plumages and simple songs. Two species are found in the West Indies: the blue-gray in the Bahamas, and the Cuban gnatcatcher. The family is mostly distributed in the Old World.
Cuban gnatcatcher, Polioptila lembeyei. Guantanamo Naval Base, Guantanamo, south-eastern Cuba.
(Photograph courtesy of Mr. Joseph Burgess).
Cuban gnatcatcher, Polioptila lembeyei. Playa Siboney, south-eastern Cuba.
(Photographs courtesy of Mr. Eladio M. Fernandez).
Family Tyrannidae: Tyrant Flycatchers
The tyrant flycatchers are a group of usually drab-looking birds that are generally poor songsters as well. (Indeed, the term "songbird" does not mean that a particular species within the order sings well). Kingbirds, pewees, and elaenias of one sort or another inhabit every Caribbean island. Notorious for being rather pugnacious (and hence the name of the entire family), kingbirds are creatures of savannas, woodlands, and forest canopies. Elaenias and pewees are smaller, and occupy similar general habitats, but are segregated from kingbirds and each other by occupying somewhat different structural niches, especially in forests' understories and in scrubland. While most species are insectivores, larger ones will capture and devour prey like lizards frogs.
Some kingbirds are so aggressive as to give chase and harass far larger birds like hawks and herons whenever these enter their territories. Only hummingbirds in this region are as bold in attacking intruders.
Loggerhead kingbird (Puerto Rican race, Tyrannus caudifasciatus taylori).
First photograph: El Yunque National Forest, north-eastern Puerto Rico.
Second photograph: Maricao State Forest, western Puerto Rico.
This species utters a loud and rasping call, especially in the morning.
Loggerhead kingbird (Jamaican race, Tyrannus caudifasciatus jamaicensis). Windsor, north-central Jamaica.
Its rolling and rather soft call is most often heard in the early morning.
Gray kingbirds, Tyrannus dominicensis. Eggleston, south-central Dominica, Lesser Antilles.
Known for their pugnacious attitude, members of Tyrannus will not hesitate to attack hawks, herons, and other large birds that enter into their territory.
A gray kingbird (Greater Antillean race, Tyrannus dominicensis dominicensis) captures a anthophorid bee to feed its chicks.
This is one of the most widespread birds in the Caribbean, and its call is heard in almost every island.
Cabo Rojo State Forest, south-western Puerto Rico.
Gray kingbird, Tyrannus dominicensis dominicensis, showing the orange patch on its crown in an aggressive display.
Carolina, north-eastern Puerto Rico.
Gray kingbird nest. Guanica State Forest, south-western Puerto Rico.
Caribbean elaenia (northern race, Elaenia
martinica riisii).
First photograph: Lajas, south-western Puerto Rico.
Second
photograph: Guanica State Forest, south-western Puerto Rico.
This species inhabits the Lesser Antilles and reaches the Greater
Antilles in the Virgin Islands and Puerto Rico.
A male Caribbean elaenia emits its call from a tree top.
Eggleston, south-central Dominica, Lesser Antilles.
Greater Antillean elaenia, Elaenia fallax. Blue Mountains, east-central Jamaica.
Puerto Rican pewee, Contopus portoricensis.
First photograph: Maricao State Forest, western Puerto Rico.
Second photograph: Susua State Forest, south-western Puerto Rico.
Its sweet and plangent call resembles the sound of water filling a crystal goblet.
Hispaniolan pewee, Contopus hispaniolensis. Puerto Escondido, south-western Dominican Republic, Hispaniola.
Crescent-eyed pewees, Contopus caribaeus. Cueva de Los Peces, south-western Cuba.
The species is also found in the Bahamas.
(Photograph courtesy of Mr. Eladio M. Fernandez).
Lesser
Antillean pewee, Contopus latirostris. Syndicate,
north-central Dominica, Lesser Antilles.
Saint Lucian pewee, Contopus oberi.
Grande Anse, north-eastern Saint Lucia, Lesser Antilles.
La Sagra's flycathcher, Myiarchus sagrae. Bermejas, south-western Cuba.
(Photograph courtesy of Mr. Eladio M. Fernandez).
Stolid flycatcher, Myiarchus stolidus. Near Rabo de Gato, south-western Dominican Republic, Hispaniola.
(Second photograph courtesy of Mr. Pedro Genaro Rodriguez).
Puerto Rican flycatchers, Myiarchus antillarum.
First two photographs: Lajas, south-western Puerto Rico.
Last photograph: Guanica State Forest, south-western Puerto Rico.
Lesser Antillean flycatcher, Myiarchus oberi. Syndicate, north-central Dominica, Lesser Antilles.
Grenadian flycatcher, Myiarchus nugator.
This species is endemic to the insular banks of Saint Vincent and Grenada.
First photograph: Mount Hartman Bay, south-western Grenada, Lesser Antilles.
Second photograph: Kingstown Botanical Garden, south-western Saint Vincent, Saint Vincent and the Grenadines, Lesser Antilles.
Rufous-tailed flycatcher, Myiarchus validus. Blue Mountains, east-central Jamaica.
This Jamaican endemic is the largest member of its genus in the Caribbean.
And this, also a Jamaican endemic, is the smallest of its genus in the Caribbean: the sad Flycatcher, Myiarchus barbirostris.
First photograph: Windsor, north-central Jamaica.
Second photograph: Blue Mountains, east-central Jamaica.
Its call is softer than those of other members of the genus.
Family Vireonidae: Vireos
Vireos are small birds similar to wood warblers, but with heavier bills and of more phlegmatic disposition that is in accordance with feeding more heavily on fruits than on insects. Their melodious but somewhat monotonous calls are heard in most wooded areas in the Antilles.
Each of the Greater Antilles has one or two endemic species.
Jamaican vireos, Vireo jamaicensis. Blue Mountains, east-central Jamaica.
Black-whiskered vireo, Vireo altiloquus. Blue Mountains, east-central Jamaica.
The call of this species is one of the most commonly heard sounds in the West Indies, in summertime.
Flat-billed vireo, Vireo nanus. Puerto Escondido, Bahoruco Mountains, south-western Hispaniola.
This species is endemic to its island.
(Photograph courtesy of Mr. Miguel Angel Landestoy).
Cuban vireo, Vireo gundlachii. Las Salinas, south-western Cuba.
(Photograph courtesy of Mr. Eladio M. Fernandez).
Thick-billed vireo, Vireo crassirostris. Providenciales, Turks and Caicos.
(Photograph courtesy of Mr. Joseph Burgess).
Family Corvidae: Crows and Their Kin
This group includes crows, ravens, magpies, and jays. Only crows have reached the West Indies, where there are several endemic species in the Greater Antilles, including some that have sadly become extinct in historic times. Most members of the family lack overly melodious voices, and their repertoires are limited to screams, honks, and other simple notes. However, some Antillean emit series of irregular and liquid notes that persons used to Holartic species find very unfamiliar.
Fabled for being among the most cunning and resourceful beings with feathers, the biggest crows (ravens) are also the largest members of their order.
Palm crow, Corvus palmarum. Enriquillo National Park, south-western Dominican Republic, Hispaniola.
Endemic to Cuba and Hispaniola.
Jamaican crow, Corvus jamaicensis. Windsor, north-central Jamaica.
As with other Caribbean species, its calls include some un-crow-like notes.
Cuban crow, Corvus nasicus. Varadero, south-western Cuba.
(Photograph courtesy of Mr. Eladio M. Fernandez).
White-necked crow, Corvus leucognaphalus. Los Haitises National Park, northern Dominican Republic, Hispaniola.
This impressive bird is named after the white bases of its neck feathers, not seen unless the animals puffs up its plumage.
Presently restricted to Hispaniola, this largest corvid in the West Indies formerly inhabited Puerto Rico, as well.
Its voice is rather unusual for a crow, and consists of a gurgling series of melodious notes mixed with harsher ones.
(Photographs courtesy of Mr. Pedro Genaro Rodriguez).
Family Hirundinidae: Swallows
The Cosmopolitan swallows are the best known and most easily recognized members of the immense order of songbirds, as well as the most aerial among them. Morphologically and ecologically similar to the unrelated swifts, West Indian swallows often make their mud nests under roof overhangs and bridges, as well as on cliffs and caves. There are several endemic species and subspecies in the Bahamas and the Greater Antilles.
A Caribbean Martin chick begs food from a somewhat unwilling mother. Piñones, north-eastern Puerto Rico.
(Photograph courtesy of Dr. Luis O. Nieves).
Barn swallow, Hirundo rustica, juvenile. Grand Anse, south-western Grenada, Lesser Antilles.
This species, widespread in the Northern Hemisphere winters in the West Indies in considerable numbers.
Golden swallow, Tachycineta euchrysea. Aceitillar, Bahoruco Mountains, south-western Dominican Republic, Hispaniola.
This species is endemic to Hispaniola and Jamaica.
(Photograph courtesy of Mr. Miguel Angel Landestoy).
Family Troglodytidae: Wrens
Wrens are generally small and drably-colored birds, yet many have powerful and melodious voices. In several of the Lesser Antilles, the tiny Troglodytes aedon emits its hyperbolic song from the branches of the rainforest's understory, and it even inhabits some suburban areas, in gardens and yards. This bird also lives in North America, from where vagrants sometimes arrive in the Bahamas. Some taxonomists consider the Lesser Antillean populations to be a different species.
The only other genus that inhabits the West Indies is monotypic. Larger than the house wren, the Zapata wren, Ferminia cerverai, is endemic to a small area in south-western Cuba.
House wren (Dominican race, Troglodytes aedon rufescens). A tiny bird with an immense voice.
It's beautiful, if jumbled, song announces its presence even as the bird is absurdly small and not easily seen.
Eggleston, south-central Dominica, Lesser Antilles.
Family Turdidae: Thrushes, Solitaires, and Their Kin
This family is represented in the West Indies by the widespread Turdus thrushes and the Myadestes solitaires, and by Cichlherminia lherminieri (belonging to a monotypic Lesser Antillean genus). Some of these species emit beautiful and melodious songs, heard especially in the rain and humid forests of the islands.
As well, there are several migrants of this family, belonging to diverse genera, to be found in the Antilles.
Red-legged thrush (Hispaniolan and Puerto Rican race, Turdus plumbeus plumbeus).
First photograph: Carolina, north-eastern Puerto Rico.
Second photograph: Enrique Marti Coll Park, San Juan, north-eastern Puerto Rico.
This species is restricted to the Greater Antilles, except for one population inhabiting the Lesser Antillean island of Dominica.
Its sweet, if monotonous, song is often heard in both cities and rural areas before dawn and in late afternoons.
When alarmed, it emits a scolding "wet-wet" call.
It is one of the more terrestrial species of the genus in the Caribbean, often seen seeking insects and other invertebrates in leaf litter.
The Cuban race of the red-legged thrush has a reddish tint on its lower abdomen.
Cueva de Los Peces, south-western Cuba.
(Photograph courtesy of Mr. Eladio M. Fernandez).
White-chinned thrush, Turdus aurantius. Anchovy, north-western Jamaica.
This species ecologically substitutes the previous one in its own island.
La Selle's thrush, Turdus swalesi. Zapoten, south-western Dominican Republic, Hispaniola.
Endemic to Hispaniola, this is a species of montane rain and pine forests.
(Photograph courtesy of Mr. Pedro Genaro Rodriguez).
Bared-eyed robin, Turdus nudigenis. Kingstown Botanical Garden, south-western Saint Vincent, Saint Vincent and the Grenadines, Lesser Antilles.
This species reaches through the Lesser Antilles north to Martinique, and is widespread in South America.
Rufous-throated solitaire, Myadestes genibarbis. Blue Mountains, east-central Jamaica.
The haunting song of this species consists of prolonged, high-pitched whistles with a strange "electronic" quality.
It may also emit a trill at the end of a phrase.
It indeed resembles a sweetly melodious version of the noise produced by a microphone with "feedback".
The song is extremely ventriloqual and far-carrying, beginning and fading at inaudible levels,
so it can be very difficult to locate the bird by its voice.
Conversely, the alarm or aggressive note is a harsh and piercing "pe-oh".
(Audio file courtesy of Dr. Luis O. Nieves).
Rufous-throated solitaire, Myadestes genibarbis. Cortico, Bahoruco Mountains, south-western Dominican Republic, Hispaniola.
(Photograph courtesy of Mr. Miguel Angel Landestoy).
Rufous-throated solitaire, (Dominican race, Myadestes genibarbis dominicanus). Syndicate, north-central Dominica, Lesser Antilles.
Family Mimidae: Mockingbirds, Thrashers, Tremblers, and Their Kin
Thrashers, mockingbirds, and their allies are conspicuous birds found both in forest and savannas, as well as cities. Some have the capacity to imitate the songs of other birds, hence the family's name ("mimics"). The several species of Mimus mockingbirds present in the region are among the best Antillean songsters and are surpassed, on that regard, only by some of the Turdus and Myadestes thrushes.
Allenia, Margarops, and Ciclocerthia thrashers are endemic to the Lesser Antilles, one species (Margarops fuscatus) having reached Puerto Rico quite recently. Where it becomes accustomed to people, the pearly-eyed thrasher will use every given opportunity to steal some food from human tables. It also has something of a bad reputation for devouring every bird's nestling that it can kill.
The two species of Cinclocerthia are called "tremblers", and have earned their name due to the peculiar habit of quivering their wings when they are excited for any reason.
Several
species of
mimid thrushes (Mimidae) inhabit the Caribbean.
This is a pearly-eyed thrasher
(nominate race, Margarops
fuscatus fuscatus).
Guanica State Forest, south-western Puerto Rico.
A Lesser Antillean
species that invaded the Puerto Rican bank at the beginning of the
twentieth
century.
Its call somewhat resembles the tuning up of a radio.
(Audio file courtesy of Dr. Luis O. Nieves).
Scaly-breasted thrashers, Allenia fusca. Eggleston, south-central Dominica, Lesser Antilles.
Brown trembler, Cinclocerthia ruficauda. Northern Forest Reserve, Dominica, Lesser Antilles.
The name derives from its habit of vibrating the wings when it is alarmed.
Perhaps the most familiar of the mimid thrushes, and one of the most cherished birds in North America and the Greater Antilles:
the northern mockingbird, Mimus polyglottos (this one belonging to the subspecies orpheus).
Both its generic name ("imitator") as well as its specific epithet ("many-tongued") allude to its capacity
to integrate the calls and songs of other birds into its own repertoire. In the Caribbean, one often thinks he is
hearing the call of a vireo, warbler, thrush, or tanager, when actually it is a mockingbird which is being heard.
Carolina, north-eastern Puerto Rico.
A northern mockingbird, Mimus polglottos orpheus, momentarily
regurgitates a fruit in order to accommodate it better in its crop.
Carolina, north-eastern Puerto Rico.
A tropical mockingbird, Mimus gilvus, extricates its meal of insect larvae from a ripe guava, Psidium guajava.
Eggleston, south-central Dominica, Lesser Antilles.
Bahamian mockingbird, Mimus gundlachii. Cabo Rojo Natural Wildlife Reserve, south-western Puerto Rico.
This individual was found very far away from its usual range. I probably was a stray due to one of the tropical storms of the region.
(Photograph courtesy of Dr. Luis O. Nieves).
Family Dulidae: Palm Chat
The monotypic family Dulidae is probably related to waxwings. The palm chat obtains its name from its habit of building their large communal nests among palm fronds, most often. The family is endemic to Hispaniola alone, in the Greater Antilles.
Palm chats, Dulus dominicus. Santa Cruz de Barahona, south-western Dominican Republic, Hispaniola.
This monotypic family is confined to this island.
Palm chat chick. Santo Domingo, southern Dominican Republic, Hispaniola.
(Photograph courtesy of Mr. Miguel Angel Landestoy).
The communal nests of palm chats can be seen
on the crowns of trees.
Near Rabo de Gato, south-central Dominican Republic, Hispaniola.
Parulidae: Wood Warblers
The group containing the wood warblers is especially species-rich in the Antilles. Active insect-eaters, the majority of the species find their prey on trees (although a few are mainly terrestrial). They jump, flit, and fly up and down the foliage at a frantic pace, and seldom stay put for more than a few seconds except to utter their songs during mating seasons.
Many are winter migrants in the region but many others are native, even endemic, to the Caribbean islands. Some species are highly specialized, like Saint Lucia's Semper's warbler. Others are partial to particular habitats, like the Puerto Rican elfin woods warbler and the Jamaican arrowhead warbler, two sister species that inhabit montane rain forests in their respective islands.
Above and below: yellow warblers, (Lesser Antillean race, Dendroica petechia petechia), males. Eggleston, south-central Dominica, Lesser Antilles.
This is one of the most widespread birds of the Western Hemisphere, breeding from Alaska to northern South America.
It is very probable that this taxon is actually a complex of several related species which could be split apart in the near future.
The colors of a juvenile yellow warbler are more subdued than the adult's.
Eggleston, south-central Dominica, Lesser Antilles.
Yellow warbler, (Puerto Rican and Virgin Islands race, Dendroica petechia cruciana), male. Boqueron State Forest, south-western Puerto Rico.
Yellow warbler, (Puerto Rican and Virgin Islands race, Dendroica petechia cruciana), female, feeding on cactus fruit.
Boqueron State Forest, south-western Puerto Rico.
Adelaide's warblers,
Dendroica adelaidae, males. Guanica State Forest, south-western
Puerto Rico.
A Puerto Rican endemic
whose closest relatives are in Barbuda and Saint Lucia, Lesser Antilles.
Its call is a ascending trill.
(Audio file courtesy of Dr. Luis O. Nieves).
Saint Lucian warbler, Dendroica
delicata, male. Grande Anse, north-eastern Saint Lucia, Lesser
Antilles.
Together with the Adelaide's (above) and Barbudan warblers, it
comprises a superspecies.
Arrowhead warbler, Dendroica pharetra. Blue and John Crow Mountains National Park, east central Jamaica.
This form forms a triad of species together with the plumbeous warbler (below) and the elfin woods warbler of Puerto Rico.
Elfin woods warbler, Dendroica angelae. Maricao State Forest, western Puerto Rico.
A juvenile plumbeous warbler (yellowish individual) Dendroica plumbea, begs food from an adult, while another forages alone for food.
First photograph: Northern Forest Reserve, Dominica, Lesser Antilles.
Second photograph: Eggleston, south-central Dominica, Lesser Antilles.
The species is endemic to that island and Guadeloupe.
The relationships of some Caribbean species of warbler are obscure, and perhaps they should be placed in other families, like that of tanagers. These include the white-winged (Xenoligea montana) and green-tailed warbler (Microligea palustris) of Hispaniola, and the two Teretristis warblers of Cuba
Green-tailed warbler, Microligea palustris. Puerto Alejandro, southwestern Dominican Republic, Hispaniola.
This species is of uncertain relations, and could possibly be a tanager.
(Photograph courtesy of Mr. Pedro Genaro Rodriguez).
White-winged warbler, Xenoligea montana. Zapoten, south-western Dominican Republic, Hispaniola.
(Photograph courtesy of Mr. Miguel Angel Landestoy).
Yellow-headed warbler, Teretristis fernandinae. Las Salinas, south-western Cuba.
(Photograph courtesy of Mr. Eladio M. Fernandez).
Oriente warbler, Teretristis fornsi. Siboney, south-western Cuba.
(Photograph courtesy of Mr. Eladio M. Fernandez).
The West Indies are either stopovers or final destinations for a large number of migrant parulines which nest in North America. On their way south, a number of species use the Antilles as supply stations where they feed and rest, before continuing onwards to Central and South America. Other migrants remain here during the late fall to early spring, adding to the numbers of other birds found in the Antilles during those seasons. Some North American migrant birds, like the northern parula (Parula americana) and several wood warblers of the genus Dendroica winter almost exclusively in the West Indies.
The next photographs show some of the North American emberizids that either pass through or stay in the Antilles during the northern hemisphere's fall, winter, and early spring.
Black-throated blue warbler, Dendroica caerulescens, male. Carite State Forest, east-central Puerto Rico.
Black-throated blue warbler, Dendroica caerulescens, female.
Blue Mountains, east-central Jamaica.
American redstart, Setophaga ruticilla. Blue Mountains, east-central Jamaica.
Northern parula, Parula americana, male. Cambalache State Forest, northern Puerto Rico.
This individual already exhibits its breeding plumage before returning to its breeding grounds in North America.
Black-and-white warbler, Mniotilta varia. Parque Central, San Juan, north-eastern Puerto Rico.
This is among the most common northern migrants in the Antilles.
(Photograph courtesy of Dr. Luis O. Nieves).
Coerebidae: Honeycreeper
The bananaquit is the single representative of its family. This birds specialize in feeding on nectar, but also consume large quantities of small invertebrates. In fact, the species has been placed at different times among tanagers and wood warblers, though most authors place it in its own family.
This is perhaps the most ecologically versatile bird in the Caribbean islands, since it is found in every habitat that contains at least some trees. In many forests it is the most abundant bird, and it has quite a number of color morphs, showing bright yellow bellies and gray throats in the Greater Antilles, white throats in the Bahamas, and almost totally black plumages in some of the Lesser Antilles.
Bananaquit, (Puerto Rican race, Coereba flaveola portoricensis). Carolina, north-eastern Puerto Rico.
This tanager-like species feeds mainly on nectar and small insects.
Its partiality to sweets makes it easy for people in the Caribbean to attract them by the dozen to gardens and verandas, by use of a simple bowl filled with sugar.
This is one of the most widespread birds of the American tropics, found from extreme southern Florida to Argentina.
It also represents a biogeographic oddity in the Caribbean, in that it is absent from mainland Cuba, while it
inhabits every other island from the northernmost Bahamas to Grenada, southernmost of the Lesser Antilles.
A bananaquit (Virgin Islands race, Coereba flaveola sancti-thomae) feeds on the pulp and seeds of the fruit of a Pilosocereus royenii cactus.
Little Dix Bay, Virgin Gorda, British Virgin Islands.
Bananaquit feeding on ripe bananas. Eggleston, south-central Dominica, Lesser Antilles.
Bananaquit. Southern Andros Island. Bahamas.
Many Bahamian populations of these bird have white throats and flanks.
(Photograph courtesy of Mr. Joseph Burgess).
The melanistic morph of the bananquit exists in some of the southern Lesser Antilles.
First photograph: Mount Hartman Bay, south-western Grenada, Lesser Antilles.
Second photograph: Grand Anse, south-western Grenada, Lesser Antilles.
A pair of bananaquits (each of a different morph) feed their nestlings.
In the third photograph, the parent takes the excrement of one of the chicks away from the nest.
Grand Anse, south-western Grenada, Lesser Antilles.
Bananaquits have a habit of building nests not just for raising young, but for sleeping at night, as well.
Guilarte State Forest, central Puerto Rico.
Thraupidae: Tanagers
Tanagers are mainly herbivores. They feed mostly on fruits, flower buds, and leaves, though many species include varied amounts of insects in their diets. These last include the three Calyptophilus chat tanagers of Hispaniola as well as the Puerto Rican tanager, Nesospingus speculiferus.
The fact is that leaves and buds are not very good food. They are difficult to digest, and are but a meager source of energy. Such is the reason why birds and other endotherms that feed mainly on such items spend a lot of time eating.
Without a doubt, the most beautiful West Indian members of this group are the four Spindalis striped-headed tanagers of the Greater Antilles and the Bahamas. Females are always a drab olive or grayish brown with yellowish tinges, but males are striking in their bright yellow and orange bodies and black and white heads, wings, and tails.
Puerto Rican striped-headed tanager, Spindalis portoricensis, male in aggressive display.
Carolina, north-eastern Puerto Rico.
The genus Spindalis is a near-endemic to the West Indies, since the four species are mainly Greater Antillean and Bahamian in their distribution.
Outside the Caribbean region the Western stripe-headed tanager, of Cuba and the Bahamas,
has also invaded the continental island of Cozumel, off the eastern coast of Mexico.
Hispaniolan striped-headed tanager, Spindalis dominicensis, male. Bahoruco National Park, south-western Dominican Republic, Hispaniola.
Jamaican striped-headed tanager, Spindalis nigricephalus, male and female, feeding on flower buds and leaves
(Rubus sp., in the case of the male; Persea sp., in the case of the female).
Blue and John Crow Mountains National Park, east central Jamaica.
Other Antillean tanagers include the two palm tanagers of the genus Phaenicophilus. Both are Hispaniolan endemics.
Black-crowned palm tanagers, Phaenicophilus palmarum.
First photograph: National Botanical Garden, southern Dominican Republic, Hispaniola.
Second photograph: Ebano Verde Nature Reserve, Central Mountain Range, Dominican Republic, Hispaniola.
The second individual is feeding on a guava, Psidium guajava.
The two-species genus is endemic to that island.
(Photographs courtesy of Mr. Pedro Genaro Rodriguez).
Puerto Rican tanagers, Nesospingus speculiferus. El Yunque National Forest, north-eastern Puerto Rico.
Its call is a series of harsh "chek" notes.
Western chat tanager, Calyptophilus tertius. Zapoten, Bahoruco Mountains, southwestern Dominican Republic, Hispaniola.
(Photograph courtesy of Mr. Miguel Angel Landestoy).
Family Emberizidae: Bullfinches, Grassquits, and Their Kin
This group is composed of seed-eating birds with short, conical, powerful bills. Although most are rather somber in their colors, several species, like some Loxigilla bullfinches, emit loud and melodious songs easily heard amongst the foliage of their forested haunts. Other species most common in open and grassy areas, like Tiaris and Ammodramus possess thin and reedy voices.
Puerto Rican bullfinch, Loxigilla portoricensis, adult and juvenile. Maricao State Forest, western Puerto Rico.
Its loud song consisting of rising, rich whistles, is heard in most of the island's forests.
Occasionally it emits a "koochi-koochi-koochi..." call, as well.
The genus Loxigilla, composed of four species, is an exclusively Antillean endemic.
A juvenile Puerto Rican bullfinch feeds on a sphynxid moth caterpillar. Jose Luis Monagas Park,, Bayamon, north-eastern Puerto Rico.
Puerto Rican bullfinch, male. Cartagena Lagoon National Wildlife Refuge, south-western Puerto Rico.
The striking black and red coloration is actually difficult to distinguish among the dense canopy and shrubbery where it lives.
(Photograph courtesy of Dr. Luis O. Nieves).
Greater Antillean bullfinch, Loxigilla violacea, female. Bahoruco National Park, south-western Dominican Republic, Hispaniola.
A male Lesser Antillean bullfinch, Loxigilla noctis. Morne Trois Pitons National Park, central Dominica, Lesser Antilles
Unlike the case of the previous species, this taxon is sexually dichromatic except for
populations in the island of Barbados, where both sexes are olive brown in color.
Lesser Antillean bullfinch, male. Grand Anse, south-western Grenada, Lesser Antilles.
Much bolder than its Greater Antillean congeners, the Lesser Antillean bullfinch frequently raids tables in search of food scraps left by humans.
This male carries nest material. Kingstown, south-western Saint Vincent, Saint Vincent and the Grenadines, Lesser Antilles.
A female Lesser Antillean bullfinch pulls fibers from a rope, to be used in building its nest.
Eggleston, south-central Dominica, Lesser Antilles.
Cuban bullfinch, (mainland Cuban race, Melopyrrha nigra nigra). La Gran Piedra, south-western Cuba.
(Photograph courtesy of Mr. Eladio M. Fernandez).
A sub-adult male black-faced grassquit (Puerto Rican race, Tiaris bicolor omisus) patrols its territory.
Carolina, north-eastern Puerto Rico.
Black-faced grassquit (Puerto Rican race, Tiaris bicolor omisus), male.
Guanica State Forest, south-western Puerto Rico.
A black-faced grassquit (Puerto Rican race, Tiaris bicolor omisus)
interrupts its meal of buttonwood mangrove fruits to declare its territory by song.
Parque Central, San Juan, north-eastern Puerto Rico.
Black-faced grassquits, (Jamaican race, Tiaris bicolor marchii), male and female.
First photograph: Anchovy, north-western Jamaica.
Second photograph: Windsor, north-central Jamaica.
A black-faced grassquit, female, feeds on seeds.
Eggleston, south-central Dominica, Lesser Antilles.
Yellow-faced grassquit, (Puerto Rican race, Tiaris olivaceus bryanti), two males and a female
Cartagena Lagoon National Wildlife-Refuge, south-western Puerto Rico.
Yellow-faced grassquit, (Jamaican race, Tiaris olivaceus olivaceus), male and female.
Anchovy, north-western Jamaica.
Yellow-shouldered grassquit, Loxipasser anoxanthus.
Blue and John Crow Mountains National Park, east-central Jamaica.
Grasshopper sparrow (Puerto Rican race, Ammodramus savannarum boriquensis).
Caño Tiburones Wildlife Reserve, Arecibo, northern Puerto Rico.
(Photograph courtesy of Dr. Luis O. Nieves).
Orangequits, Euneornis campestris. First two photographs: male and female; Blue Mountains, east-central Jamaica.
Last two photographs: males; Anchovy, north-western Jamaica.
Although it behaves much like a honeycreeper, this bird seems to be a very specialized and derived bullfinch.
Rufous-collared sparrow, Zonotrichia capensis. Near Constanza, central Dominican Republic, Hispaniola.
This species has a peculiar distribution, being found in Central and South America but, in the Caribbean, only in the mountains of Hispaniola.
(Photograph courtesy of Mr. Miguel Angel Landestoy).
Cardinalidae: Cardinals, Saltators, and Their Kin
Saltators have powerful bills adapted to feed on seeds and, to a lesser extent, on fruits. The sole West Indian species emits a loud, rich song consisting of piercing whistles. Indeed, their calls are among the most characteristic diurnal sounds in Lesser Antillean forests.
Lesser Antillean saltator, Saltator albicollis, feeding on a papaya fruit. Cabrits National Park, north-western Dominica, Lesser Antilles.
Saltators are very large finches whose loud, whistled songs are heard in some of the Lesser Antilles in most habitats except the highest mountain peaks.
Family Icteridae: Grackles, Cowbirds, Blackbirds, Orioles, and Their Kin
Grackles, orioles, blackbirds and their allies are rather large birds with long, pointed bills adapted for feeding on insects as well as fruits. Almost all Caribbean species have some amount of glossy black on their plumage. However, many species are also strikingly marked with yellow, orange, or red.
Carib grackles, Quiscalus lugubris, displaying male and two females.
Grand Anse, south-western Grenada Lesser Antilles.
Being to a great degree onmivorous, Lesser Antillean grackles feed on anything from food scraps dropped by people, to lizards.
Grand Anse, south-western Grenada, Lesser Antilles.
Greater Antillean grackles (Puerto Rican race, Quiscalus niger brachypterus).
First two photographs: male and Female, Enrique Marti Coll Park, San Juan, north-eastern Puerto Rico.
Next two photographs: female and a brownish juvenile, Carolina, north-eastern Puerto Rico.
Similar in appearance to a grackle, this female shiny cowbird, Molothrus bonariensis, is a nest parasite.
Frequently seen in mixed flocks with their relatives, grackles and blackbirds, cowbirds have seriously impaired
the reproductive success of some endangered avian species in the Antilles.
Maricao State Forest, western Puerto Rico.
Cuban blackbirds, Dives atroviolacea. Guantanamo Naval Base, Guantanamo, south-eastern Cuba.
(Photograph courtesy of Mr. Joseph Burgess).
Yellow-shouldered blackbirds (mainland Puerto Rican race, Agelaius xanthomus xanthomus).
Cabo Rojo State Forest, south-western Puerto Rico.
Formerly widespread in most littoral areas of the island, and ranging inland on a seasonal basis, this bird is now seriously endangered
due to habitat destruction, compounded more recently by nest-parasitism by the glossy cowbird, an invader from the Lesser Antilles.
Another subspecies inhabits small Mona Island, west of Puerto Rico.
Troupial (northern South American race, Icterus icterus ridgwayi). Lajas, south-western Puerto Rico.
This beautiful bird is found in Puerto Rico and the Virgin Islands and, occasionally, in some of the Lesser Antilles.
It is not known for certain if it is native in the West Indies, or if it was introduced from South America.
Its name derives from its loud call.
Puerto Rican oriole, Icterus portoricensis. Maricao State Forest, western Puerto Rico.
(Photograph courtesy of Dr. Luis O. Nieves).
The nest of the Puerto Rican oriole, like those of its relatives, is a pendent structure that is sown on the leaves of palms and other trees.
Here an adult brings an anole to feed the chicks. These however, seem to actually be those of the parasitic glossy cowbird, Molothrus bonariensis.
Often this bird will utter its lovely song for only a few minutes before sunrise. It does the same seldom during the day, when it mostly emits a contact call.
Maricao State Forest, western Puerto Rico.
Hispaniolan oriole, Icterus dominicensis. Enriquillo National Park, south-western Dominican Republic, Hispaniola.
Jamaican oriole (Jamaican race, Icterus leucopteryx leucopteryx). Anchovy, north-eastern Jamaica.
Its sweet calls are heard in many of the rural areas of the island.
Eastern meadowlark, Sturnella magna. Playa Siboney, Santiago de Cuba, south-western Cuba.
(Photograph courtesy of Mr. Eladio M. Fernandez).
Family Fringillidae: Siskins, Euphonias, and Their Kin
Siskins (subfamily Carduelinae) and euphonias (subfamily Euphoniinae) are very small birds that feed mostly on fruits and seeds. Their beautiful colors stand out amidst the foliage of the forests they inhabit. Some species, like the Antillean euphonia, Euphonia musica, are highly specialized in feeding almost exclusively on the fruits of but a few species of plants, like mistletoes and figs.
Antillean euphonia, (Puerto Rican race, Euphonia musica sclateri), male, feeding on Ficus fruits.
Maricao State Forest, western Puerto Rico.
Formerly placed among tanagers, the several members of the genus Euphonia are now placed by taxonomists among the cardueline finches.
These tiny but brightly colored bird has a specialized diet composed mostly of mistletoe fruits.
The species has an extensive range covering many of the Lesser Antilles and the Greater Antilles save for Jamaica,
where a related endemic species lives. Lesser Antillean males resemble females in their more somber olive green color.
(Photograph courtesy of Dr. Luis O. Nieves).
Antillean euphonia, (Puerto Rican race, Euphonia musica sclateri), juvenile female.
Guanica State Forest, south-western Puerto Rico.
Antillean euphonia chicks throw themselves out of the nest, and are then fed on the ground by their parents until they are able to fly.
The adults regurgitate fruits directly into the chicks throats.
Maricao State Forest, western Puerto Rico.
Jamaican euphonias, Euphonia jamaica, males.
Blue Mountains, east central Jamaica.
Hiapniolan corssbills, Loxia megaplaga. Bahoruco National Park, south-western Dominican Republic, Hispaniola.
These finches have assymetric bills adapted to probe into pine cones and extract their seeds.
Hispaniolan siskin, Carduelis dominicensis.
Bahoruco National Park, south-western Dominican Republic, Hispaniola.